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由肌纤维膜开始，包围肌原纤维至肌浆网的细的横断小管。是兴奋收缩关连的媒介，呈平坦袋状，长径 15— 50纤米，短径 3— 5纤米。在两个肌浆网间有三分体（ triad）结构，包围着肌原纤维，如果从肌原纤维的轴以直角作成断面，则呈网眼状，总起来称 T系统（ T system）。肌纤维兴奋发生活动电位时，它传给直接开口于膜的 T管，到达肌浆网，放出 Ca。如果将肌肉用甘油温和地处理一下，结果只有 T管被破坏，膜虽发生活动电位，但肌肉不收缩。这是瑞典的安德森塞德格林（ E Andersson Cecle rgren， 1951）最先观察到的。
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