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文献和实验指DNA双链的局部，由具有互补性单链 DNA与之结合所产生的环状结构。当 DNA复制开始，在原来的双链中仅一方被新合成的短 DNA单链被置换的情况下可以见到（为 Displacement loop之简称）。可通过人工使 DNA单链结合来制成此结构。由 RNA单链所产生的类似结构称为 R环。
Analysis of Geminivirus DNA Replication by 2-D Gel
The technique described was developed for the separation of begomovirus DNA. DNA products resulting from and during geminiviral replication are characterized by the application of strand-specific separation and identification by strand
2-D Gel Electrophoresis: Constructing 2D-Gel Proteome Reference Maps
isoelectric focusing, which separates proteins according to their isoelectric point (pI), and SDS-PAGE, which separates them further according to their molecular mass. An evolution of conventional 2-DE is represented by the 2D-Difference in Gel Electrophoresis
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