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羽芽 feather bud， feather germ 鸟类羽毛的原基。其发生的第一阶段是由于真皮的肥厚而形成表皮突起，称为羽乳头（ featherpapilla）。羽乳头生长，顶端朝后方。真皮肥厚而构成其轴即羽髓（ feather pulp）。血管进入其中，表皮将其覆盖。围绕在羽髓基部的表皮从皮肤表面陷入，形成羽囊（ feather follicle）。被覆羽髓的表皮增殖，不久以羽髓为中心形成柱状体，以后角化而变成羽干，其顶端分枝而生出刷子状的羽枝。羽干的基部在真皮深层形成羽根。
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